Western Ecuador Moist Forests
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The Western Ecuador Moist Forests (NT0178), also known as th
Pacific Forest of Ecuador
is an ecoregion in the plains and western foothills of the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
of southern Colombia and Ecuador. At one time this region contained dense forests with highly diverse flora and fauna, and many endemic species. Most of the original habitat has now been destroyed, and the ecoregion is one of the most threatened in the world.


Geography


Location

The Western Ecuador moist forests ecoregion covers land to the west of the Andes in Ecuador and southern Colombia. In the northwest and the extreme southwest the ecoregion extends to the Pacific Ocean. The ecoregion is bounded on the north by the
Patía River The Patía River () is a river in southwestern Colombia. It flows over to drain into the Pacific Ocean north of Tumaco. The Patía River is the longest river on the Colombian Pacific Coast. The last is navigable by boat. Geography The Patía ...
. It extends through the provinces of Esmeraldas, Manabí and Guayas, reaching the Gulf of Guayaquil in the south, and the foothills on the Andes in the east. It varies in width from . It has an area of . In the extreme north the ecoregion transitions into the
Chocó–Darién moist forests The Chocó–Darién moist forests (NT0115) is an ecoregion in the west of Colombia and east of Panama. The region has extremely high rainfall, and the forests hold great biodiversity. The northern and southern parts of the ecoregion have been cons ...
ecoregion. To the east it transitions into the Northwestern Andean montane forests. Along the Pacific coast in some areas there are sections of South American Pacific mangroves, and in others there are sections of Ecuadorian dry forests. In the south the ecoregion merges into Guayaquil flooded grasslands.


Terrain

The ecoregion covers the coastal plain and reaches into the foothills of the Andes to the east up to an elevation of about . The region has had strong volcanic activity, with volcanic rocks found throughout the region, alternating with marine sediments from the
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
era (65 million to 2.6 million years ago). The soils are fertile, particularly in the north-central area, which covers volcanic ash and
lapilli Lapilli is a size classification of tephra, which is material that falls out of the air during a volcanic eruption or during some meteorite impacts. ''Lapilli'' (singular: ''lapillus'') is Latin for "little stones". By definition lapilli range ...
.


Climate

The ecoregion receives high rainfall with no significant dry season. Average annual rainfall is highest in the north, with over and in some areas over . In the south annual rainfall averages , and in all areas exceeds . Average annual temperatures vary from , with little change throughout the year. At a sample location at coordinates the
Köppen climate classification The Köppen climate classification is one of the most widely used climate classification systems. It was first published by German-Russian climatologist Wladimir Köppen (1846–1940) in 1884, with several later modifications by Köppen, nota ...
is Am (equatorial; monsoonal). Mean temperatures range from in July to in April. Total yearly rainfall is about . Monthly rainfall ranges from in August to in February.


Ecology

The ecoregion is in the
neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
realm, in the
tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discon ...
biome. The forests have many endemic species of plants and animals found only locally on mountain ridges or narrow strips of land.


Flora

Natural fragmentation of the western moist forests has resulted in rapid evolution of new endemic species. Around 10,000 species of plants have been reported, of which about 2,500 are endemic. The forest has a dense canopy that exceeds in height, and has many
liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a t ...
s and
epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
s, including many endemic species. Epiphytes of the
Araceae The Araceae are a family of monocotyledonous flowering plants in which flowers are borne on a type of inflorescence called a spadix. The spadix is usually accompanied by, and sometimes partially enclosed in, a spathe (or leaf-like bract). A ...
and
Cyclanthaceae Cyclanthaceae is a family of flowering plants. Taxonomy Earlier systems, such as the Cronquist system and the Takhtajan system, placed it as the sole family in the order Cyclanthales. In the classification system of Dahlgren the Cyclanthaceae w ...
families are abundant on the lower trunks of the trees. There are also mosses, lichens, ferns and palms in the dense understory. 1,250 species of plants in 136 families have been found in a single area of . According to a 1990 report, 122 new orchid species had recently been identified. 43 of the species are endemic. Emergent species, which may exceed in height, include the strangler fig ('' Ficus dugandii''). Large, dominant canopy trees include ''
Brosimum utile ''Brosimum utile'' (Kunth), also called ''Galactodendron'', Pittier (= ''B. galactodendron)'' is a plant species in the family Moraceae. Description ''Brosimum utile'' can grow to a height of 30m. It is monoecious and has bisexual inflorescence ...
'', ''
Carapa guianensis ''Carapa guianensis'' is a species of tree in the family Meliaceae, also known by the common names andiroba or crabwood. Description Andiroba is native to the Amazon and is widely used by the indigenous populations of the northern region o ...
'', '' Guarea kunthiana'' and '' Virola dixonii''. Other trees include ''
Clarisia racemosa ''Clarisia'' is a genus of trees in the family (biology), family Moraceae, native to North America, North and South America (from Mexico to Bolivia). Taxonomy The genus ''Clarisia'' contains the following species: * ''Clarisia biflora'' Ruiz & ...
'', '' Matisia coloradorum'', '' Pourouma chocoana'', ''
Pouteria ''Pouteria'' is a genus of flowering trees in the gutta-percha family, Sapotaceae. The genus is widespread throughout the tropical regions of the world. It includes the canistel ('' P. campechiana''), the mamey sapote ('' P. sapota''), and the l ...
'' species, '' Pseudolmedia eggersii'' and ''
Symphonia globulifera ''Symphonia globulifera'', commonly known as boarwood, is a timber tree abundant in Central America, the Caribbean, South America and Africa. This plant is also used as a medicinal plant and ornamental plant. Common names Common trade names of t ...
''. The endemic tree '' Humiriastrum procerum'' grows to the north of the Guayllabamba River, and is often logged for use in construction. Common understory species include the ''
Iriartea deltoidea ''Iriartea'' is a genus in the palm family Arecaceae, native to Central and South America. The best-known species – and probably the only one – is ''Iriartea deltoidea'', which is found from Nicaragua, south into Bolivia and a great p ...
'' and ''
Wettinia quinaria ''Wettinia'' is a palm genus, consisting of flowering plants in the family Arecaceae. The genus, established in 1837, contains some 20 species, but more seem to await discovery considering that 4 species - '' W. aequatorialis'', '' W. lanata'', ...
'' palms. Rare species such as '' Dicliptera dodsonii'' are found at the Río Palenque Scientific center in Esmeraldas. There are records of rare or extinct flora such as ''
Carapa megistocarpa ''Carapa megistocarpa'' is a species of flowering plant in the family Meliaceae. It is endemic to Ecuador, where it is a tree of coastal forests. It is endangered by deforestation Deforestation or forest clearance is the removal of a fo ...
'' and '' Erythrochiton carinatus''.


Fauna

According to a 1990 report, 5 new mammal species had recently been identified. The
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
(''Panthera onca'') is endangered. Other endangered mammals include
Baird's tapir The Baird's tapir (''Tapirus bairdii''), also known as the Central American tapir, is a species of tapir native to Mexico, Central America, and northwestern South America. It is the largest of the three species of tapir native to the Americas, as ...
(''Tapirus bairdii''),
equatorial dog-faced bat The equatorial dog-faced bat (''Cabreramops aequatorianus'') is a species of bat in the family Molossidae. It is endemic to Ecuador. They are found in dry, tropical forests. The species is now endangered. The equatorial dog-faced bat feeds on in ...
(''Molossops aequatorianus'') and Geoffroy's spider monkey (''Ateles geoffroyi''). Endangered reptiles include Boulenger's least gecko ('' Sphaerodactylus scapularis''), green sea turtle (''Chelonia mydas'') and
hawksbill sea turtle The hawksbill sea turtle (''Eretmochelys imbricata'') is a critically endangered sea turtle belonging to the family Cheloniidae. It is the only extant species in the genus ''Eretmochelys''. The species has a global distribution, that is largel ...
(''Eretmochelys imbricata''). The
American crocodile The American crocodile (''Crocodylus acutus'') is a species of crocodilian found in the Neotropics. It is the most widespread of the four extant species of crocodiles from the Americas, with populations present from South Florida and the coasts ...
(''Crocodylus acutus'') is also threatened. 650 species of birds were identified in a 1993 survey, of which many are threatened. Species found in the most humid areas include the Baudo guan (''Penelope ortoni''),
Berlepsch's tinamou Berlepsch's tinamou (''Crypturellus berlepschi'') is a type of ground bird found in moist forestBirdLife International (2008) in northwestern Colombia and northwestern Ecuador.Clements, J (2007) Taxonomy The Berlepsch's tinamou is a monotypic s ...
(''Crypturellus berlepschi''),
blue-whiskered tanager The blue-whiskered tanager (''Tangara johannae'') is a species of bird in the family Thraupidae. It is found in the Chocó of Colombia and Ecuador. Its natural habitat is subtropical or tropical moist lowland forests. It is threatened by habitat ...
(''Tangara johannae''), brown wood rail (''Aramides wolfi''),
crowned woodnymph The crowned woodnymph or violet-crowned woodnymph (''Thalurania colombica'') is a species of hummingbird in the "emeralds", tribe Trochilini of subfamily Trochilinae. It is found from Belize and Guatemala to northern Peru.HBW and BirdLife Int ...
(''Thalurania colombica''),
El Oro parakeet The El Oro parakeet, conure D'Orcès, cotorra de El Oro, or perico de El Oro (''Pyrrhura orcesi'') is a species of parrot in the family Psittacidae, endemic to Ecuador. It is a relatively newly identified species, having been discovered in 1980. L ...
(''Pyrrhura orcesi''), El Oro tapaculo (''Scytalopus robbinsi''), indigo flowerpiercer (''Diglossa indigotica''), long-wattled umbrellabird (''Cephalopterus penduliger'') and scarlet-breasted dacnis (''Dacnis berlepschi''). Species found in less humid areas include
grey-backed hawk The grey-backed hawk (''Pseudastur occidentalis'') is an Endangered species of bird of prey in subfamily Accipitrinae, the "true" hawks, of family Accipitridae. It is found in Ecuador and far northern Peru.Remsen, J. V., Jr., J. I. Areta, E. Bo ...
(''Pseudastur occidentalis''), ochraceous attila (''Attila torridus'') and rufous-headed chachalaca (''Ortalis erythroptera'').
Humboldt's sapphire Humboldt's sapphire or Humboldt's hummingbird (''Chrysuronia humboldtii'') is a species of hummingbird in the "emeralds", tribe Trochilini of subfamily Trochilinae. It is found in Colombia, Ecuador, and Panama. Taxonomy and systematics Humbol ...
(''Hylocharis humboldtii'') is found only in the mangroves to the north. Species that have almost been wiped out due to hunting and
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological processes ...
include
crested guan The crested guan (''Penelope purpurascens'') is a member of an ancient group of birds of the family Cracidae, which are related to the Australasian megapodes or mound builders (Megapodiidae). It is found in the Neotropics, in lowlands forests ran ...
(''Penelope purpurascens''), great curassow (''Crax rubra''),
great green macaw The great green macaw (''Ara ambiguus''), also known as Buffon's macaw or the great military macaw, is a Central and South American parrot found in Nicaragua, Honduras, Costa Rica, Panama, Colombia and Ecuador. Two allopatric subspecies are rec ...
(''Ara ambiguus''),
great tinamou The great tinamou (''Tinamus major'') is a species of tinamou ground bird native to Central and South America. There are several subspecies, mostly differentiated by their coloration. Taxonomy The great tinamou was described and illustrated in 1 ...
(''Tinamus major'') and
harpy eagle The harpy eagle (''Harpia harpyja'') is a neotropical species of eagle. It is also called the American harpy eagle to distinguish it from the Papuan eagle, which is sometimes known as the New Guinea harpy eagle or Papuan harpy eagle. It is the ...
(''Harpia harpyja''). Endangered birds include
banded ground cuckoo The banded ground cuckoo (''Neomorphus radiolosus'') is a Endangered species of cuckoo in the tribe Neomorphini of subfamily Crotophaginae. It is found in Colombia and Ecuador.HBW and BirdLife International (2021) Handbook of the Birds of the ...
(''Neomorphus radiolosus''), Baudo guan (''Penelope ortoni''),
Esmeraldas woodstar The Esmeraldas woodstar (''Chaetocercus berlepschi'') is a rare, neotropical species of hummingbird in the family Trochilidae. There are six different species in the woodstar genus''.'' Most of them are poorly studied due to their small size, ext ...
(''Chaetocercus berlepschi''),
great green macaw The great green macaw (''Ara ambiguus''), also known as Buffon's macaw or the great military macaw, is a Central and South American parrot found in Nicaragua, Honduras, Costa Rica, Panama, Colombia and Ecuador. Two allopatric subspecies are rec ...
(''Ara ambiguus''),
grey-backed hawk The grey-backed hawk (''Pseudastur occidentalis'') is an Endangered species of bird of prey in subfamily Accipitrinae, the "true" hawks, of family Accipitridae. It is found in Ecuador and far northern Peru.Remsen, J. V., Jr., J. I. Areta, E. Bo ...
(''Pseudastur occidentalis''),
grey-cheeked parakeet The grey-cheeked parakeet (''Brotogeris pyrrhoptera''), less commonly known as fire-winged parakeet, is a species of parrot in the family Psittacidae. Taxonomy The grey-cheeked parakeet was described and named by John Latham in 1801. Descript ...
(''Brotogeris pyrrhoptera''), rufous-brown solitaire (''Cichlopsis leucogenys'') and
slaty becard The slaty becard (''Pachyramphus spodiurus'') is a species of bird in the family Tityridae. It has traditionally been placed in Cotingidae or Tyrannidae, but evidence strongly suggest it is better placed in Tityridae,Atelopus balios ''Atelopus balios'', the Rio Pescado stubfoot toad, is a species of toad in the family Bufonidae. It is endemic to southwestern Ecuador, with records from Pacific lowlands in Azuay, Cañar, and Guayas Provinces. It is a rare species that was ...
''), elegant stubfoot toad (''Atelopus elegans''),
phantasmal poison frog The phantasmal poison frog or phantasmal poison-arrow frog (''Epipedobates tricolor'') is a species of poison dart frog. It is endemic to Ecuador and known from the Andean slopes of central Ecuador in Bolívar Province. They have radiant colors ...
(''Epipedobates tricolor''), horned marsupial frog (''Gastrotheca cornuta''), Pichincha rocket frog ('' Hyloxalus toachi''), '' Pristimantis colomai'', spring robber frog ('' Pristimantis crenunguis''), Alto Tambo rain frog ('' Pristimantis degener''), hotel robber frog ('' Pristimantis tenebrionis''), blue-spotted toad (''
Rhaebo caeruleostictus ''Rhaebo caeruleostictus'' is a species of toad in the family Bufonidae. It is endemic to Ecuador and occurs along the lower western slope of the Cordillera Occidental at elevations of asl. The specific name ''caeruleostictus'', from Latin ' ...
'') and Rio Pitzara robber frog ('' Strabomantis helonotus'').


Status

The
World Wide Fund for Nature The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wor ...
(WWF) gives the region the status of "Critical/Endangered". Construction of highways and exploration for oil between 1960 and 1980 caused the destruction of most of the moist forests, now one of the most threatened of the world's habitats. Ongoing main threats come from banana plantations and palm oil and rubber extraction. Most of the moist forests in the ecoregion are in the Esmeraldas province between the San Lorenzo Canton south of the Chocó forests of Colombia and Quinindé. There are only small remnants of the original forest elsewhere in the region, and these are degraded near their margins. There are fairly well preserved narrow strips of forest parallel to the mountains in the north, but these are being damaged by settlements and indiscriminate logging. There is some protection in the Cotacachi Cayapas Ecological Reserve, and the highlands from are within the
Machalilla National Park Machalilla National Park ( es, Parque Nacional Machalilla) is a preserve in the Ecuadorian Manabí Province near Puerto López and the rural parish of Machalilla, a small fishing village in the vicinity of the park. Established in 1979, Machal ...
. The Rio Palenque Scientific Station contributes to research and protection.


References


Bibliography

* * * {{Biodiversity of Colombia Ecoregions of Colombia Ecoregions of Ecuador Neotropical tropical and subtropical moist broadleaf forests